Morphology of callus, shoots, roots and leaves of Withania somnifera (Cultivated and Wild) in vitro tissue culture conditions with different hormones concentration

Modern plant tissue culture is practiced under aseptic or favourable conditions under filtered air. Living plant materials which was taken from the environment or free space are being naturally contaminated or will show contamination on their external surfaces (and sometimes interiors) with pathogenic microorganisms, so surface sterilization of initial materials (explants) in chemical solutions (usually Sodium or calcium hypochlorite or mercuric chloride) is done. Mercuric chloride is mostly used as a plant sterilant now a days, unless other sterilizing agents which require for removal of contamination are dispose found to be least effective, as it is very dangerous and harmful to use, and is very difficult to dispose of (Aniel, et al., 2011) [1]. 

Explants are then generally placed or maintained on the upper surface of a solidified culture medium (normally MS) (Pawar & Maheshwari, 2004) [11], but during some cases it will be inoculate directly into a liquid nutrient medium, normally when cell suspension cultures are required. Solid nutrient and liquid nutrient media usually consists of hormones. Solidified media are prepared or produced from the liquid media with the use of a gelling or solidified agent, generally a purified agar. The constituents of the
solid medium or the requirement of the plant growth hormones and the nitrogenous source (nitrate/ammonium salts and amino acids) have produces morphological effects of the grown tissues that from the initial explants. For example, high concentration of auxin responsible for the high multiplication of roots whereas an excess cytokinin may yield shoots. 

A balance concentration of auxin and cytokinin will often generate an unorganised and undifferentiated growth of cells, i.e. callus (Sabir, et al., 2008) [14], but the morphology of the outgrowth depends on the different plant varieties as well as the suitable medium composition (Joshi & Padhya, 2010) [5]. As cultures grow, explants transferred to new media (sub-cultured) to allow for growth or to change the morphology of the culture. Gita Rani and Avinash, (2003) [4] observed or reported the callus induction or cultivation fromthe hypocotyls, root and cotyledonary leaf segments of Ashwagandha grown on MS medium provided with various different concentrations and combinations of the 2,4-dichlorophenoxyacetic acid (2,4-D) and Kinetin plant growth harmone. Maximum callusing (near about 100%) was observed with root and cotyledonary leaf explants segments grown on Murashige and Skoog medium provided with a combination of 2,4-di-chlorophenoxyacetic acid (2,4-D) and Kinetin plant growth harmone.

Gita Rani and Avinash (2003) [4] also told that or suggested that the frequency of the formation of root was comparatively lower or become slow with the higher concentrations of Indole 3- butyric acid (IBA) either alone or in combination with indole 3- acetic acid (IAA) or NAA and with all concentrations of indole 3- acetic acid (IAA) alone in Ashwagandha. 


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